Veröffentlicht in Sauropod Vertebra Picture of the Week

What does it cost to publish a paper  in a non-open access Elsevier journal? The immediate cost to the author is often zero (though page charges, and fees for colour illustrations mean this is not always true). But readers have to pay to see the paper, either directly in the case of private individuals or through library budgets in the case of university staff and students. What is the total cost to the world?